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Rev. Biol. Trop. (Int. J. Trop. Biol. ISSN-0034-7744) Vol. 64 (3): 1311-1331, September 2016
Arntz, W., Brey, E., Tarazona, J., & Robles, A. (1987).
Changes in the structure of a shallow sandy beach com-
munity in Peru during an El Niño event. South African
Journal of Marine Science, 5, 645-658.
Borja, A., Dauer, D. M., & Elliott, M. (2010). Medium and
long-term recovery of estuarine and coastal ecosys-
tems: Patterns, rates, and restoration effectiveness.
Estuaries and Coasts, 33, 1245-1260.
Brusca, R. C., Moore, W., & Shuster, S. M. (2016). Inver-
tebrates. 3
rd
edition. Sunderland, MA, USA.: Sinauer
Associates Inc.
Cameron, C. B. (2005). A phylogeny of the hemichordates
based on morphological characters. Canadian Journal
of Zoology, 83, 196-215.
Cloern, J. E., Foster, S. Q., & Kleckner, E. (2014). Phyto-
plankton primary production in the world’s estuarine-
coastal ecosystems. Biogeosciences, 11, 2477-2501.
Cutler, N., Cutler, E., & Vargas, J. A. (1992). Peanut worms
(Phylum Sipuncula) from Costa Rica. Revista de Bio-
logia Tropical,40, 153-158.
Cutler, E. B. (1994). The Sipuncula: Their systematics,
biology, and evolution. Ithaca, New York, U.S.A:
Comstock Publishing Association.
Dall, W. H. (1921). Annotated list of Recent Brachiopoda in
the collections of the United States National Museum,
with descriptions of thirty-three new forms. Procee-
dings of the U. S. National Museum, 57, 261-377.
Deland, C., Cameron, C. B., Rao, K. P., & Bullock, T. H.
(2010). A taxonomic revision of the family Harri-
maniidae (Hemichordata: Enteropneusta) with des-
cription of seven species from the Eastern Pacific.
Zootaxa, 2408, 1-30.
De Moreno, J. E. A., Gerpe, M. S., Moreno, V. S., & Vodo-
pivez, C. (1997). Heavy metals in Antartic organisms.
Polar Biology, 17, 131-140.
Dittmann, S., & Vargas, J. A. (2001). Tropical tidal flat
benthos compared between Australia and Central Ame-
rica. In K. Reise (Ed.), Ecological Comparisons of
Sedimentary Shores (pp. 275-293). Vol. 151, Ecologi-
cal Studies. Berlin, Germany: Springer.
Dittmann, S. (2002). Benthic fauna in tropical tidal flats - a
comparative perspective. Wetlands Ecology and Mana-
gement, 10, 189-195.
Emig, C. C., & Vargas, J. A. (1990). Glottidia audebarti
(Broderip) (Brachiopoda, Lingulidae) from the Gulf
of Nicoya, Costa Rica. Revista de Biología Tropical,
38, 251-258.
Emig, C. C. (1983). Taxonomie du genre Glottidia (Bra-
chiopodes, Inarticulés). Bulletin du Muséum National
d’Histoire Naturelle, Paris. 4
e
Ser. 5. Section A. n
o
.
2, 469-489.
Emig, C. C. (1997). Ecology of inarticulate brachiopods. In
R. L. Kaesler (Ed.), Treatise on Invertebrate Paleon-
tology, Part H. Brachiopoda revised (pp. 473-495).
Boulder, Colorado & Laurence, Kansas. U.S.A.: Geo-
logical Society of America and University of Kansas.
Emig, C. C. (2009). Brachiopods. Text: pp. 417-420.
Species list: CD pp. 119-121. In I. S. Wehrtmann &
J. Cortés (Eds.). Marine Biodiversity of Costa Rica,
Central America. Monogr. Biol. 86. Berlín: Springer
+ Business Media B.V.
García-Céspedes, J., Acuña-González, J., & Vargas-Zamora,
J. A. (2004). Metales traza en sedimentos costeros de
Costa Rica. Revista de Biología Tropical, 52 (Supl.
2), 51-60.
García, V., J., Acuña-González, J., Vargas-Zamora, J. A.,
& García-Céspedes, J. (2006). Calidad bacterioló-
gica y desechos sólidos en ambientes costeros de
Costa Rica. Revista de Biología Tropical, 54 (Supl.
1), 35-48.
Gray, J. S., & Elliott, M. (2009). Ecology of Marine Sedi-
ments- from science to management. 2
nd
ed. Oxford,
U.K.: Oxford University Press.
Gravel, P., Johanning, K., McLachlan, J., Vargas, J. A., &
Oberdorster, E. (2006). Imposex in the intertidal snail
Thais brevidentata (Gastropoda-Muricidae) from the
Pacific coast of Costa Rica. Revista de Biología Tropi-
cal, 54 (Suppl. 1), 21-26.
Ha, N. T. T., Nhuan, M. T., Ngoc, N. T., & Dung, H. T.
(2007). The distribution of peanut-worm (Sipunculus
nudus) in relation to geo-environmental characteristics.
VNU Journal of Sciences, Earth Sciences, 23, 110-115.
Hunter, C. L., Krane, E., & Fitzpatrick, J. (2008). Current
and historic distribution and abundance of the inarti-
culate brachiopod, Lingula reevii Davidson (1880) in
Kaneohe Bay, Oahu, Hawaii, U.S.A. Marine Biology,
155, 205-210.
Hyman, L. H. (1959). The Invertebrates. Vol. V: Smaller
Coelomate Groups – Chaetognatha, Hemichordata,
Pogonophora, Phoronida, Ectoprocta, Brachiopo-
da, Sipunculida. The coelomate bilateria. New York,
U.S.A.: McGraw-Hill Book Company Inc.
Jones, G. F., & Barnard, J. L. (1963). The distribution and
abundance of the inarticulate btachiopod Glottidia
albida (Hinds) on the mainland shelf of Southern
California. Pacific Naturalist, 4, 27-52.
Jope, H. M. (1965). Composition of the brachiopod shell.
In R. C. Moore (Ed.), Treatise on Invertebrate
Paleontology, Vol. I. Part. H. Brachiopoda (pp. 156-
164). Kansas, U.S.A.: The Geological Society of
America-The University of Kansas Press.
Kawauchi, G. Y., & Giribet, G. (2013). Sipunculus nudus
Linnaeus, 1766 (Sipuncula): cosmopolitan or a
group of pseudo-cryptic species? Marine Ecology,
2013, 1-14.