Mendelson III, J R, E D Brodie, J H Malone, M E Acevedo, M A Baker, N J Smatresk, and J A Campbell. 2004. “Factors Associated With the Catastrophic Decline of a Cloudforest Frog Fauna in Guatemala”. Revista De Biología Tropical 52 (4):991–1000. https://revistas.ucr.ac.cr/index.php/rbt/article/view/15596.