Heyer, W. Ronald. 1968. “Studies on the Genus <i>Leptodactylus</i> (Amphibia: Leptodactylidae). II. Diagnosis and Distribution of the <i>Leptodactylus</i> Of Costa Rica”. Revista De Biología Tropical 16 (2):171–205. https://doi.org/10.15517/rev.biol.trop. v16i2.28149.