Wille, Alvaro. 1973. “Coleoptera: Scarabaeidae Observations on the Behavior of a Tropical Rain Forest Dung Beetle, <i>Megathoposoma candezei</i≫ (Harold)”. Revista De Biología Tropical 21 (1):41–57. https://revistas.ucr.ac.cr/index.php/rbt/article/view/27322.