Travis, B. V., M. Vargas V., and J. C. Swartzwelder. “Bionomics of Black Flies (Diptera: Simuliidae) in Costa Rica. I. Species Biting Man, With an Epidemiological Summary for the Western Hemisphere”. Revista De Biología Tropical, vol. 22, no. 1, July 1974, pp. 187–200, https://revistas.ucr.ac.cr/index.php/rbt/article/view/26090.