Wille, Alvaro. “Phylogenetic Significance of an Unusual African Stingless Bee, Meliponula Bocandei (Spinola)”. Revista de Biología Tropical 11, no. 1 (July 1, 1963): 25–45. Accessed January 11, 2025. https://revistas.ucr.ac.cr/index.php/rbt/article/view/31135.