Lankesteriana: International Journal on Orchidology
type: channel
description: LANKESTERIANA, International Journal on Orchidology, is an international scientific journal published by the Lankester Botanical Garden Research Center – University of Costa Rica. The journal is globally recognized for its specialization and contributions to the knowledge of the orchid family. In addition to research articles, LANKESTERIANA is dedicated to facilitating scholarly communication and exchange by publishing a comprehensive array of content, including book reviews, checklists, comments, conference proceedings, correspondence, corrigenda, obituaries, opinions, reviews, and special issue contributions. LANKESTERIANA maintains rigorous academic standards as a peer-reviewed journal, ensuring that only the highest quality research articles are published. Through this rigorous review process, the journal maintains its reputation for excellence and reliability, providing readers with access to credible, authoritative scientific information on orchids. Furthermore, LANKESTERIANA values contributions from authors worldwide, with English as the official language of the journal. Through Early View, LANKESTERIANA is committed to reducing the time to effective publication of scientific, peer-reviewed papers, for the benefit of both authors and the journal's readers. The papers in this section of Lankesteriana receive an individual DOI and are published online as soon as the editorial process is concluded positively. They represent early views of the final papers exactly as they will appear in the hard-printed, complete issue of the journal. The effective date of publication of each paper is established by the date of its first publication online, which is recorded on the front page of the paper, both in its online and hard-printed versions. The printed version of LANKESTERIANA is published periodically in volumes by the Lankester Botanical Garden, University of Costa Rica. Each volume is made up of three issues published in April (covers the period from January to Apr…
publisher: Universidad de Costa Rica
language: en
publication name: Lankesteriana: International Journal on Orchidology
issn: 1409-3871
copyright: According to the Open Access policy promoted by the University of Costa Rica, all the papers published by Lankesteriana are licensed under the Creative Commons copyright and can be downloaded free of charge. The journal holds copyright and publishing rights under the CC BY-NC-ND 3.0 CR license. Before the publication of the materials submitted by the author(s) in LANKESTERIANA, the author(s) hereby assign all rights in the article to the Lankester Botanical Garden.
items: Female orchid bee Euglossa dilemma visits the perfume orchid Coryanthes panamensis in Florida
items: A new species of Catasetum (Cymbidieae: Catasetinae) from the Colombian Amazonia
items: Catasetum tavaresii (Catasetinae), a new species from Central Brazilian Amazon
items: The fragrant orchid Vanilla chamissonis: an appraisal of the genetic structure of wild populations from Osununú Natural Reserve (Misiones, Argentina)
items: Additions to orchid flora of Bhutan III: Paphiopedilum × pradhanii
items: Floral visitors of three species of Corallorhiza in Monte Tláloc, Texcoco, Estado De México, Mexico
items: new species of Brachionidium (Pleurothallinidae) from the Yanachaga Chemillen National Park, Pasco, Peru
items: new endemic species of Polystachya (Polystachyinae) from eastern Tanzania
items: Orchids of the Southern Cone (1830–2000) – Part IV. Maevia Noemí Correa’s Flora Patagonica
items: Telipogon pillaropatatensis (Oncidiinae): a new species from the East-Central Andes of Ecuador
items: Sarcoglottis wernerherzogii (Spiranthinae), a new species from Cusco, Peru
items: Odontoglossum crispum: a tale of love, loss and scientific discovery
items: Studies on Costa Rican Vanilloideae: the return of Cleistes rosea and a reaffirmation of Epistephium ellipticum
items: Notes on the correct identity of Ceratostylis siamensis (Epidendroideae: Podochileae) from India
items: Lepanthes chalalensis (Pleurothallidinae), a new species endemic to the Santander department in Colombia
items: New species and records of Orchidaceae from Costa Rica. IV.
items: Bulbophyllum lampadion (section Monanthes), a new species from New Guinea
items: A new species of Dendrobium sect. Crinifera (Dendrobieae) from Thailand
items: Corybas geminigibbus, a new addition to the orchid flora of Sumatra
items: Addition of five orchid species to the flora of Bhutan
items: Mormodes benelliana (Catasetinae), a new species from Mato Grosso, Brazil
items: new green-flowered Aa (Cranichideae) from Peru
items: new species Telipogon (Oncidiinae) from Bolivia
items: Rostellum in orchids
items: Ten new orchid records from the Indonesian archipelago
items: Orchids naturalized in Florida
items: The genus Encyclia Hook. (Orchidaceae, Epidendroideae) in Maranhão state, Brazil: new morphological data and extent of distribution
items: More than a century later: first modern record of Pteroglossa macrantha (Orchidaceae: Spiranthinae) in Argentina, with notes on its morphology, distribution, habitat, and conservation implications
items: A new species of Pleurothallis (Pleurothallidinae) from the P. talpinaria complex, with notes on comparative floral morphology and pollination ecology
items: Addition of Five New Species to the Orchid Flora of Bhutan IV

Female orchid bee Euglossa dilemma visits the perfume orchid Coryanthes panamensis in Florida
type: item
description: Male orchid bees gather fragrance chemicals from the surfaces of about 700 species of neotropical orchids and pollinate them in the process. The males display the collected chemicals during their courtship, and female orchid bees will mate only with males, displaying a species-specific blend of collected chemicals. Given this reality, the discovery of a female Euglossa dilemma dead in the bucket trap of a flower of the perfume orchid Coryanthes panamensis bearing the orchid’s pollinarium is extraordinary. There are no previous reports of female orchid bees visiting a perfume orchid and picking up pollinia of the orchids. Bearing a pollinarium indicates that this female bee made at least two visits to the flowers and that these visits were unlikely to be accidental. The flower in which the bee was found was on a cultivated plant grown in a residential garden in Coral Gables, Florida. Neither the orchid nor the orchid bee are native to Florida and occur in different areas of tropical America. Why this female orchid bee visited this orchid is considered, including the possibility that it may be a gynandromorph. Despite being a morphological female, the bee may have had neurological anomalies that caused it to exhibit a male-like attraction to this perfume orchid.
subject: Correspondence
  topic: section
subject: bucket orchid
  topic: keywords
subject: fragrance chemicals
  topic: keywords
subject: gynandromorphism
  topic: keywords
subject: naturalized
  topic: keywords
creator: Robert W. Pemberton
rights: Copyright (c) 2026 Lankesteriana: International Journal on Orchidology https://creativecommons.org/licenses/by-nc-nd/3.0/cr/
date: 2026-05-19
publication date: 2026-05-19
starting page: 277
ending page: 283
doi: 10.15517/lank.v24i3.61983

A new species of Catasetum (Cymbidieae: Catasetinae) from the Colombian Amazonia
type: item
description: A new species, Catasetum caquetense from the Caquetá department of the Colombian Amazonia, is described and illustrated. Catasetum caquetense is morphologically distinct from similar species such as C. tuberculatum in its rectangular, entire, denticulate, prominent apiculate basal callus surrounded by inconspicuous crown-shaped protrusions with an extra callus towards the apex that partially narrows towards the mid-lip. Information about its distribution, ecology, habitat, and taxonomic comments on morphologically similar species is also provided. With this addition, the genus comprises eight species in the department of Caquetá.
subject: Articles
  topic: section
subject: Amazonia
  topic: keywords
subject: Catasetum caquetense
  topic: keywords
subject: Catasetum tuberculatum
  topic: keywords
subject: orchid
  topic: keywords
subject: Caquetá
  topic: keywords
creator: R. Alexis Calderón Álvarez
creator: Miguel M. Bonilla Morales
rights: Copyright (c) 2023 Lankesteriana: International Journal on Orchidology https://creativecommons.org/licenses/by-nc-nd/3.0/cr/
date: 2026-05-13
publication date: 2026-05-13
starting page: 469
ending page: 475
doi: 10.15517/lank.v23i3.54462

Catasetum tavaresii (Catasetinae), a new species from Central Brazilian Amazon
type: item
description: A new Catasetum species from Brazilian Amazon, more specifically from the “Floresta Nacional de Saracá-Taquera” (State of Pará), is proposed, described and illustrated. This new taxon belongs to the subgenus Catasetum, section Isoceras, subsection Isoceras, whose representatives have symmetrical and parallel antennae. It is compared with C. nhamundaense, C. garnettianum and C. barbatum, all with the same antennae characteristics and belonging to the "C. cristatum alliance". Among related species, C. tavaresii differs mainly in the lip structure. We also give data on geographical distribution, habitat, phenology, flower visitors, and preliminary conservation status.
subject: Articles
  topic: section
subject: biodiversity
  topic: keywords
subject: Catasetum cristatum alliance
  topic: keywords
subject: epiphyte
  topic: keywords
subject: Neotropics
  topic: keywords
subject: State of Pará
  topic: keywords
subject: Taxonomy
  topic: keywords
creator: Dayse Raiane Passos Krahl
creator: Miguel Sena de Oliveira
creator: Philippe Schmal
creator: Guy Chiron
creator: João Batista Fernandes da Silva
creator: Amauri Herbert Krahl
creator: Sheylla Susan Moreira da Silva de Almeida
creator: Patrick de Castro Cantuária
rights: Copyright (c) 2023 Lankesteriana: International Journal on Orchidology https://creativecommons.org/licenses/by-nc-nd/3.0/cr/
date: 2026-05-13
publication date: 2026-05-13
starting page: 485
ending page: 493
doi: 10.15517/lank.v23i3.55794

The fragrant orchid Vanilla chamissonis: an appraisal of the genetic structure of wild populations from Osununú Natural Reserve (Misiones, Argentina)
type: item
description: Due to diverse factors the commercial crops of vanilla are highly vulnerable to genetic erosion. The species Vanilla chamissonis Klotzsch has been shown to produce interesting bioeconomic compounds that could diversify the market. In Argentina, the distribution of this fragrant orchid is restricted to specific north-eastern regions, like the Osununú Natural Reserve (at San Ignacio, Misiones), where it grows wildly in sectors (nuclei). This germplasm represents a locally adapted endemism, and thus is a valuable resource worth preserving. Herein, the genetic characterization of unexplored V. chamissonis´ germplasm was accomplished; as well, the specimens´ molecular identification was assessed in a phylogenetic context using nuclear and chloroplastidial markers. The genotyping exposed that, at least, four gene pools contribute to the genetic diversity of these plants, and that ca. 81% of total genetic variation is allocated within populations. Present results suggest a predominance of sexual reproduction, that population N1 emerges as a candidate unit worth conserving, and that a genetic discontinuity was evidenced between northeastern and southwestern nuclei due to their different genetic constitutions. In concurrence, a tenuous isolation by distance was also unveiled. Our phylogenetic results suggest that it is appropriate to consider the vanilla germplasm surveyed as V. chamissonis. The basic knowledge generated in this work will aid in guiding the in-situ conservation and management initiative, ongoing at the Osununú´s reserve.
subject: Articles
  topic: section
subject: molecular genotyping
  topic: keywords
subject: Misiones province
  topic: keywords
subject: Osununú
  topic: keywords
subject: phylogenetics
  topic: keywords
subject: population genetics
  topic: keywords
subject: wild orchids
  topic: keywords
creator: Jimena Cascales
creator: Carolina Paris
creator: Alexandra Gottlieb
rights: Copyright (c) 2023 Lankesteriana: International Journal on Orchidology https://creativecommons.org/licenses/by-nc-nd/3.0/cr/
date: 2026-05-13
publication date: 2026-05-13
starting page: 433
ending page: 456
doi: 10.15517/lank.v23i3.56432

Additions to orchid flora of Bhutan III: Paphiopedilum × pradhanii
type: item
description: Paphiopedilum × pradhanii, a natural hybrid between P. fairrieanum (subgenus Paphiopedilum, section Ceratopetalum) and P. venustum (subgenus Paphiopedilum, section Barbata), is reported for the first time from Bhutan. Detailed geographical distribution, ecology, and conservation status, and dichotomous keys to all Paphiopedilum species found in Bhutan are provided. A brief description of the parents of this hybrid is also provided.
subject: Articles
  topic: section
subject: Cypripedioideae
  topic: keywords
subject: natural hybrid
  topic: keywords
subject: Paphiopedilum (grex) Pandion
  topic: keywords
subject: section Barbata
  topic: keywords
subject: section Ceratopetalum
  topic: keywords
creator: Phub Dorji
creator: Phub Gyeltshen
creator: Kezang Tobgay
creator: Sherab Dorji
creator: Ugyen Dorji
creator: Sha Kinley Doya
creator: Pankaj Kumar
rights: Copyright (c) 2023 Lankester Botanical Garden, University of Costa Rica https://creativecommons.org/licenses/by-nc-nd/3.0/cr/
date: 2026-05-13
publication date: 2026-05-13
starting page: 425
ending page: 432
doi: 10.15517/lank.v23i3.57038

Floral visitors of three species of Corallorhiza in Monte Tláloc, Texcoco, Estado De México, Mexico
type: item
description: Corallorhiza is a genus of mycoheterotrophic orchids that includes 12 species, most of them restricted to North and Central America. In Mexico, there are seven taxa, distributed throughout the country, except for seven of the 32 Mexican states; in Estado de México all of them are present. The Orchidaceae is known for its diversity of pollination syndromes; however, for the more than 200 species of mycoheterotrophic orchids, little has been studied about their floral visitors and pollinators, and for the Mexican species of Corallorhiza nothing is known to date. Our goal was to document photographically and identify all the floral visitors of three sympatric species of the genus in Monte Tláloc, municipality of Texcoco, Estado de México. We observed individuals of Araneae, Coleoptera, Diptera, Hemiptera, and Hymenoptera visiting flowers of Corallorhiza macrantha, C. macrantha × C. maculata, C. maculata, and C. striata. Only Ocyptamus coeruleus and Platycheirus sp. (Diptera: Syrphidae) were found carrying pollinaria on their thorax. These are the first records of potential pollinators for the genus Corallorhiza in Mexico.
subject: Articles
  topic: section
subject: flower visitors
  topic: keywords
subject: hoverflies
  topic: keywords
subject: mycoheterotrophic plants
  topic: keywords
subject: Syrphidae
  topic: keywords
subject: terrestrial orchids
  topic: keywords
creator: Bruno E. Téllez-Baños
creator: Ivonne N. Gomez-Escamilla
creator: Alejandro Navarrete-Jiménez
creator: Adolfo Espejo-Serna
creator: Ana R. López-Ferrari
rights: Copyright (c) 2023 Lankester Botanical Garden, University of Costa Rica https://creativecommons.org/licenses/by-nc-nd/3.0/cr/
date: 2026-05-13
publication date: 2026-05-13
starting page: 457
ending page: 468
doi: 10.15517/lank.v23i3.57671

new species of Brachionidium (Pleurothallinidae) from the Yanachaga Chemillen National Park, Pasco, Peru
type: item
description: A new Brachionidium from Peru is described and illustrated based on living plants. Information on distribution, ecology, etymology, and phenology is also included. The new species, B. montieliae, was discovered in one of the highest mountains of the Oxapampa-Ashaninka-Yanesha Biosphere Reserve (BIOAY), situated above the forests of the Humid Puna of the Yanachaga Chemillen National Park at 3590 m in elevation. It is similar to Brachionidium vasquezii but differs in the oblong-elliptic, denticulate, and semi-erect leaves, somewhat lanuginose abaxially; the purple lanuginose adaxially and papular abaxially, entire, acuminate lateral sepals; the purplish oblong-ovate dorsal sepal, lanuginose on the adaxial surface, papular abaxially; the purple petals with scalloped edges, acuminate to obtuse apex ending in a thin, slightly abbreviated tail; the lip with apiculate lateral angles and with a central, trapezoid verrucose callus on the disc. Brachionidium montieliae is a terrestrial species that thrives in the humid Puna forests at elevations around 3400–3600 meters. The ecosystem is characterized by a sclerophyllous forest with a diverse range of tree species. The species typically blooms between July and August. There are potential threats from annual burning practices in the region. Given the potential risks, it is imperative to undertake immediate conservation measures and promote environmental awareness to safeguard this species.
subject: Articles
  topic: section
subject: bidentate stigma
  topic: keywords
subject: biosphere reserve Oxapampa–Ashaninka–Yanesha
  topic: keywords
subject: Brachionidium montieliae
  topic: keywords
subject: claviform pollinia (8)
  topic: keywords
subject: trapezoid callus
  topic: keywords
subject: wet puna forest
  topic: keywords
creator: Luis Valenzuela Gamarra
rights: Copyright (c) 2023 Lankester Botanical Garden, University of Costa Rica https://creativecommons.org/licenses/by-nc-nd/3.0/cr/
date: 2026-05-13
publication date: 2026-05-13
starting page: 477
ending page: 483
doi: 10.15517/lank.v23i3.58033

new endemic species of Polystachya (Polystachyinae) from eastern Tanzania
type: item
description: Polystachya nguruensis, a new endemic species from the isolated Nguru Mountains of eastern Tanzania, is described here, and its affinities and conservation status are discussed. Polystachya nguruensis differs from P. lukwangulensis in having much longer, more obviously several-noded pseudobulbs, shorter, narrowly oblong-elliptic, coriaceous leaves, and a longer, more densely many-flowered, branching inflorescence with smaller flowers and ecallose lip with a narrowly oblong a midlobe with a broadly obovate to subcircular, apiculate, lip midlobe. Unlike most allied species, it is a lithophyte, growing on sloping slabs of crystalline rock, a highly vulnerable habitat that has already been partly destroyed by fire. Consequently, it is assessed to be Critically Endangered and in need of immediate conservation attention.
subject: Invited papers
  topic: section
subject: conservation suggestions
  topic: keywords
subject: Critically Endangered
  topic: keywords
subject: endemism
  topic: keywords
subject: new species
  topic: keywords
subject: Nguru Mountains
  topic: keywords
subject: Polystachya nguruensis
  topic: keywords
subject: threats
  topic: keywords
creator: Phillip J. Cribb
creator: Andreas Hemp
rights: Copyright (c) 2023 Lankester Botanical Garden, University of Costa Rica https://creativecommons.org/licenses/by-nc-nd/3.0/cr/
date: 2026-05-13
publication date: 2026-05-13
starting page: 585
ending page: 591
doi: 10.15517/lank.v23i3.58060

Orchids of the Southern Cone (1830–2000) – Part IV. Maevia Noemí Correa’s Flora Patagonica
type: item
description: Maevia Noemí Correa (1914–2005) will always be remembered as the director and editor of an outstanding work, Flora Patagonica, in which she assumed the treatment of several plant families, among them the Orchidaceae. Published between 1969 and 1999, the work was the product of 30 years of dedicated research and fieldwork, resulting in the publication of eight volumes and extraordinary plant collections, now housed in Argentinian and foreign herbaria. Popularly known as The Dame of the Orchids, Correa must be considered one of the most relevant Argentinian botanists of the 20th century. In honor of her birth on February 14th, the day has been designated as "The Day of the Orchidophile”.
subject: Articles
  topic: section
subject: Argentina
  topic: keywords
subject: Chile
  topic: keywords
subject: Maevia N. Correa
  topic: keywords
subject: Orchidaceae
  topic: keywords
subject: Patagonia
  topic: keywords
creator: Carlos Ossenbach
rights: Copyright (c) 2023 Lankester Botanical Garden, University of Costa Rica https://creativecommons.org/licenses/by-nc-nd/3.0/cr/
date: 2026-05-13
publication date: 2026-05-13
starting page: 511
ending page: 528
doi: 10.15517/lank.v23i3.58132

Telipogon pillaropatatensis (Oncidiinae): a new species from the East-Central Andes of Ecuador
type: item
description: A new species, Telipogon pillaropatatensis, is described from the highlands of east-central Ecuador. It is morphologically similar to T. octavioi but differs by having a corolla with longitudinal thick vein lines without reticulations and a column with three dense bundles of equally sized setae. Also, our phylogenetic analyses suggest that they are not closely related. Interestingly, among the analyzed taxa, T. pillaropatatensis appears to have a closer phylogenetic relationship with the highly distinct T. pulcher and T. hausmannianus than with the morphologically similar T. octavioi. The new species has been found in a relatively small area, covering just a few square kilometers in east-central Ecuador. Additionally, we documented remarkable variation in the coloration and size of the corolla. Information regarding the species' habitat, ecology, phylogenetic relationships, and conservation status is also provided.
subject: Articles
  topic: section
subject: Llanganates National Park
  topic: keywords
subject: montane forests
  topic: keywords
subject: morphological variability
  topic: keywords
subject: Pastaza River Basin
  topic: keywords
subject: Telipogon octavioi
  topic: keywords
subject: Tungurahua Province
  topic: keywords
creator: Gabriel A. Iturralde
creator: Marco F. Monteros
creator: Marco M. Jiménez
creator: Carlos Martel
creator: Luis E. Baquero
rights: Copyright (c) 2023 Lankester Botanical Garden, University of Costa Rica https://creativecommons.org/licenses/by-nc-nd/3.0/cr/
date: 2026-05-13
publication date: 2026-05-13
starting page: 495
ending page: 509
doi: 10.15517/lank.v23i3.58100

Sarcoglottis wernerherzogii (Spiranthinae), a new species from Cusco, Peru
type: item
description: A new species of Sarcoglottis from the department of Cusco, Peru is described and illustrated as S. wernerherzogii. It is similar to eastern South American S. fasciculata but distinguished by its contrasting habitat (high-Andean montane cloud forest as 2050−2500 m a.s.l. vs. Atlantic rain forest near sea level), flowering after the shedding of the leaves (vs. flowering with leaves present); homogeneous green or yellow flowers (vs. whitish or pale green with contrasting reddish veins); rigidly conduplicate-concave lateral sepals that, when spread out are strongly falcate (vs. slightly concave, not rigid, obliquely lanceolate lateral sepals); strongly cymbiform, obovate hypochile as wide as long when spread out (vs. broadly channeled, oblanceolate hypochile about twice longer than wide or longer when spread out); ovate, strongly deflexed epichile provided with a central longitudinal thickening throughout its length (vs. erect or gently arcuate epichile with a thickening only at the apex); and apical anther beak truncate (vs. apical anther beak rounded). The strongly cymbiform hypochile that, when spread out is as wide as long, is unique in the genus. The differences between S. fasciculata, the new species, and other Peruvian species of Sarcoglottis are contrasted in a table. Sarcoglottis wernerherzogii is known from less than a dozen plants in two populations, both under nominal protection in the Machupicchu Historical Sanctuary and the Area of Conservación Regional Choquequirao, respectively. The status “DD” (data deficient) is suggested until an objective assessment of its risk status can be carried out.
subject: Invited papers
  topic: section
subject: Eastern Andean Cordillera
  topic: keywords
subject: Machupicchu Historical Sanctuary
  topic: keywords
subject: Sarcoglottis fasciculata
  topic: keywords
creator: Benjamín Collantes
creator: José D. Edquén
creator: Feliciano Incahuamán
creator: Gerardo A. Salazar
rights: Copyright (c) 2023 Lankester Botanical Garden, University of Costa Rica https://creativecommons.org/licenses/by-nc-nd/3.0/cr/
date: 2026-05-13
publication date: 2026-05-13
starting page: 623
ending page: 632
doi: 10.15517/lank.v23i3.58135

Odontoglossum crispum: a tale of love, loss and scientific discovery
type: item
description: This is the story of Odontoglossum crispum (Oncidium alexandrae), from the 19th century to the present day. It is a love story because Odontoglossum crispum was considered by many to be the most beautiful of all orchid species. It was collected by the hundreds of thousands from the cloud forests of its native Colombia and exported to Britain, the rest of Europe and the USA. Vast numbers were lost in transit. When they arrived many perished due to a lack of understanding of its needs in cultivation. Only the ‘best’ forms were grown, and the remainder discarded. Today much of its native habitat has either disappeared or been despoiled due to human actions. Few plants remain in cultivation in the UK. It was one of the first species to be grown commercially from seed, most notably in the UK by Charlesworth & Co., following the scientific discovery by Noël Bernard that orchid seeds required the participation of a suitable fungus to germinate under natural conditions.
subject: Invited papers
  topic: section
subject: conservation
  topic: keywords
subject: history
  topic: keywords
subject: mycorrhiza
  topic: keywords
subject: orchid hunters
  topic: keywords
subject: scrapbooks
  topic: keywords
subject: seed germination
  topic: keywords
creator: Philip T. Seaton
rights: Copyright (c) 2023 Lankester Botanical Garden, University of Costa Rica https://creativecommons.org/licenses/by-nc-nd/3.0/cr/
date: 2026-05-13
publication date: 2026-05-13
starting page: 593
ending page: 612
doi: 10.15517/lank.v23i3.58144

Studies on Costa Rican Vanilloideae: the return of Cleistes rosea and a reaffirmation of Epistephium ellipticum
type: item
description: Subfamily Vanilloideae represents a small fraction of the orchid flora of Costa Rica, and yet our knowledge of this predominantly South American group has significantly increased in the last two decades. Until recently, Vanilloideae was represented in Costa Rica by just two genera: Cleistes and Vanilla, with one and seven species, respectively. Today, up to 13 species of Vanilla are shown to occur in the country, we evidence the presence of a second species of Cleistes and confirm the presence of a third genus Vanilloideae in Costa Rica: Epistephium. The rare Cleistes rosea and Epistephium ellipticum are fully illustrated on the basis of Costa Rican materials with Lankester Composite Digital Plates (LCDP). The typification of Cleistes rosea is discussed and a color illustration is provided of C. costaricensis for comparison.
subject: Invited papers
  topic: section
subject: Cleistes costaricensis
  topic: keywords
subject: Cleistes rosea
  topic: keywords
subject: Epistephium ellipticum
  topic: keywords
subject: new country records
  topic: keywords
subject: Orchidaceae
  topic: keywords
subject: typification
  topic: keywords
creator: Adam P. Karremans
creator: Isler F. Chinchilla
creator: Gustavo Rojas-Alvarado
rights: Copyright (c) 2023 Lankester Botanical Garden, University of Costa Rica https://creativecommons.org/licenses/by-nc-nd/3.0/cr/
date: 2026-05-13
publication date: 2026-05-13
starting page: 613
ending page: 622
doi: 10.15517/lank.v23i3.58136

Notes on the correct identity of Ceratostylis siamensis (Epidendroideae: Podochileae) from India
type: item
description: While conducting taxonomic studies on the tribe Podochileae (Orchidaceae) for the Flora of India, two species, Ceratostylis siamensis and Ceratostylis himalaica have been examined using types, protologues, fresh specimens, and authentic herbarium collections. Ceratostylis siamensis was previously reported and documented in Arunachal Pradesh, India. However, we determined upon closer examination that this record was based on erroneous identification. The correct identity of this species is C. himalaica. Consequently, Ceratostylis siamensis is excluded from the flora of India.
subject: Articles
  topic: section
subject: Arunachal Pradesh
  topic: keywords
subject: Ceratostylis himalaica
  topic: keywords
subject: misidentification
  topic: keywords
subject: orchids
  topic: keywords
subject: taxonomy
  topic: keywords
creator: Shuvadip Sarkar
creator: DINESH KUMAR AGRAWALA
creator: Debabrata Maity
creator: Madhusudhan Khanal
rights: Copyright (c) 2024 Lankesteriana: International Journal on Orchidology https://creativecommons.org/licenses/by-nc-nd/3.0/cr/
date: 2026-05-13
publication date: 2026-05-13
starting page: 123
ending page: 129
doi: 10.15517/lank.v24i2.55730

Lepanthes chalalensis (Pleurothallidinae), a new species endemic to the Santander department in Colombia
type: item
description: We propose Lepanthes chalalensis as a newly identified species confined to the north-east Andes of Colombia. We randomly placed 341 sampling plots across the Eastern Cordillera, including transformed and natural habitats. Here, we provide a detailed description, illustrating images, ecological discussions, a taxonomic key for the new entity, and a conservation status analysis. The species is highly geographically restricted (in 2 out of 341 sampling plots) and has a low population size (26 adult individuals). While L. chalalensis shows a resemblance to L. velosa from Ecuador, it can be distinguished by the wide lower lobe of the petals, which is long ciliated, the lip laminae, which are reduced and bear stiff cilia, and the appendix length, which appear to be twice as long as the lip blades. The species should be considered a conservation concern due to its high rarity.
subject: Articles
  topic: section
subject: Eastern Cordillera
  topic: keywords
subject: endemic
  topic: keywords
subject: Neotropics
  topic: keywords
subject: standardised sampling
  topic: keywords
creator: Edicson Parra-Sánchez
creator: Eugenio Restrepo
creator: Yadir Barrera
creator: Juan Camilo Ordóñez-Blanco
creator: David Edwards
rights: Copyright (c) 2024 Lankesteriana: International Journal on Orchidology https://creativecommons.org/licenses/by-nc-nd/3.0/cr/
date: 2026-05-13
publication date: 2026-05-13
starting page: 131
ending page: 140
doi: 10.15517/lank.v24i2.56321

New species and records of Orchidaceae from Costa Rica. IV.
type: item
description: The fourth release of the series “New Species and Records of Orchidaceae from Costa Rica” documents and illustrates 17 taxa of the Costa Rican orchid flora. The new species records belong to the subtribes Pleurothallidinae (9 spp.), Maxillariinae (3 spp.), and Oncidiinae (2 spp.), including the description of two new species in Pleurothallis and Telipogon. Two new forms are described for Isochilus latibracteatus (Ponerinae) and Masdevallia striatella(Pleurothallidinae), and their differences from the typical forms are discussed. Additionally, the first record of a naturalized population of Phalaenopsis stuartiana(Aeridinae) in Costa Rica is discussed. Detailed descriptions, based on selected Costa Rican material, are included for all taxa, along with illustrations and/or Lankester Composite Digital Plates (LCDP). Information on their etymology, distribution, habitat, phenology, and distinguishing features compared to morphologically similar species is also provided. Finally, following the publication of the most recent Costa Rican orchid catalogue, we have identified some omissions and other new species described, which are discussed here. The orchid flora of Costa Rica now includes 1695 species and nine forms.
subject: Articles
  topic: section
subject: Aeridinae
  topic: keywords
subject: Costa Rican orchids
  topic: keywords
subject: Flora Costaricensis
  topic: keywords
subject: Maxillariinae
  topic: keywords
subject: Oncidiinae
  topic: keywords
subject: Pleurothallidinae
  topic: keywords
creator: Noelia Belfort-Oconitrillo
creator: Grettel Salguero
creator: Lizbeth Oses
creator: Karen Gil-Amaya
creator: Gustavo Rojas-Alvarado
creator: Isler F. Chinchilla
creator: Melissa Díaz-Morales
creator: Franco Pupulin
creator: Diego Bogarín
creator: Adam P. Karremans
rights: Copyright (c) 2024 Lankesteriana: International Journal on Orchidology https://creativecommons.org/licenses/by-nc-nd/3.0/cr/
date: 2026-05-13
publication date: 2026-05-13
starting page: 141
ending page: 192
doi: 10.15517/lank.v24i2.60686

Bulbophyllum lampadion (section Monanthes), a new species from New Guinea
type: item
description: Bulbophyllum lampadion from New Guinea is herein described and illustrated based on herbarium specimens and cultivated material. This high-elevation species belongs to section Monanthes, of which sect. Oxysepala is here considered to be a synonym.
subject: Articles
  topic: section
subject: hotspot
  topic: keywords
subject: Indonesia
  topic: keywords
subject: Papua New Guinea
  topic: keywords
subject: sectional classification
  topic: keywords
subject: sect. Oxysepala
  topic: keywords
creator: Jaap J. Vermeulen
creator: Malcolm Perry
creator: André Schuiteman
rights: Copyright (c) 2024 Lankesteriana: International Journal on Orchidology https://creativecommons.org/licenses/by-nc-nd/3.0/cr/
date: 2026-05-13
publication date: 2026-05-13
starting page: 193
ending page: 198
doi: 10.15517/lank.v24i2.61182

A new species of Dendrobium sect. Crinifera (Dendrobieae) from Thailand
type: item
description: A new species of Dendrobium sect. Crinifera is described from Thailand. The late Peter O’Byrne is appropriately credited for his important preparatory work for our study and for coining the name of the new species, D. krabiense. Apparently endemic to southern Thailand, D. krabiense is similar to D. pardalinum but differs in the lip mid-lobe being distinctly clawed and in the lip ornaments only consisting of two lateral keels that extend from near lip base to claw of mid-lobe. Dendrobium krabiense is also similar to D. phuketense but differs in having larger, spotted flowers with a proportionally longer mentum. The material here referred to D. krabiense was previously misidentified as Flickingeria pallens (now considered a synonym of D. pardalinum).
subject: Articles
  topic: section
subject: Dendrobium krabiense
  topic: keywords
subject: endemism
  topic: keywords
subject: Flickingeria
  topic: keywords
subject: Orchidaceae
  topic: keywords
subject: systematics
  topic: keywords
subject: taxonomy
  topic: keywords
creator: Henrik Æ. Pedersen
creator: Somran Suddee
rights: Copyright (c) 2024 Lankester Botanical Garden, University of Costa Rica https://creativecommons.org/licenses/by-nc-nd/3.0/cr/
date: 2026-05-13
publication date: 2026-05-13
starting page: 199
ending page: 203
doi: 10.15517/lank.v24i2.61571

Corybas geminigibbus, a new addition to the orchid flora of Sumatra
type: item
description: Corybas is an orchid genus of 154 species distributed from India to South China, the Western Pacific, and New Zealand. Sumatra is home to eight species, five of which are endemic. However, the true diversity is likely underestimated due to limited exploration in some parts of the island. During a recent exploration in the northern Gayo Plateau near the northern tip of the island, a Corybas species was discovered that did not match any previously known species. Further identification through specimen comparison and literature review confirmed that the species is Corybas geminigibbus. It was found growing in montane bogs. This discovery marks the second bioregion in Indonesia where this species has been recorded and the ninth Corybas species on the island. The finding also provides additional evidence of floristic similarities, suggesting past geological connections between northern Sumatra and mainland Asia. A detailed description, discussion, and photographs are provided.
subject: Articles
  topic: section
subject: monocots
  topic: keywords
subject: montane bogs
  topic: keywords
subject: taxonomy
  topic: keywords
subject: West Malesia
  topic: keywords
creator: Wendy A. Mustaqim
creator: Zulfan Arico
creator: Ridho B. Ginting
creator: Tiara Rahayu
creator: Yuda R. Yudistira
rights: Copyright (c) 2026 Lankesteriana: International Journal on Orchidology https://creativecommons.org/licenses/by-nc-nd/3.0/cr/
date: 2026-05-06
publication date: 2026-05-06
starting page: 239
ending page: 244
doi: 10.15517/lank.v24i3.60323

Addition of five orchid species to the flora of Bhutan
type: item
description: Bhutan, situated in the eastern part of the Himalaya Biodiversity Hotspot, is renowned for its rich flora and fauna. This biodiversity is attributed to the country’s multiple biogeographic origins, diverse topo- graphy, ecological complexity, and varied climatic and soil conditions. However, much of the flora remains under-collected, with many taxa yet to be discovered. From June 2022 to March 2024, floristic studies were conducted in Bhutan, leading to the discovery of five orchid species not previously recorded in the country: Bulbophyllum crabro, B. rigidum, Cymbidium tortisepalum var. longibracteatum, Galeola cathcartii, and Liparis kumokiri. These species are scientifically documented for the first time in Bhutan. Detailed descriptions, type information, updated global distribution, ecology, and colour plates of the recorded species are provided.
subject: Articles
  topic: section
subject: biodiversity
  topic: keywords
subject: distribution
  topic: keywords
subject: Orchidaceae
  topic: keywords
subject: Punakha
  topic: keywords
subject: taxonomy
  topic: keywords
creator: Phub Gyeltshen
creator: Kinley Rabgay
creator: Phuentsho
creator: Kezang Tobgay
creator: Pema Namgyel
rights: Copyright (c) 2024 Lankester Botanical Garden, University of Costa Rica https://creativecommons.org/licenses/by-nc-nd/3.0/cr/
date: 2026-05-13
publication date: 2026-05-13
starting page: 205
ending page: 216
doi: 10.15517/lank.v24i2.61573

Mormodes benelliana (Catasetinae), a new species from Mato Grosso, Brazil
type: item
description: In the present study, we describe and illustrate Mormodes benelliana, a new species found in the southwest region of the Brazilian state of Mato Grosso. This species occurs as an epiphyte in the transition between the Amazon and Cerrado domains. Mormodes benelliana is similar to M. hoehnei, but differs by having smaller pseudobulbs, inflorescences longer than the pseudobulbs, conspicuous peduncle, yellowish-brown and brown-vinous striped flowers, with a large-elliptic to ovate, glabrous lip.
subject: Articles
  topic: section
subject: Amazon
  topic: keywords
subject: Cerrado
  topic: keywords
subject: ecotone
  topic: keywords
subject: epiphyte
  topic: keywords
subject: orchid
  topic: keywords
subject: taxonomy
  topic: keywords
creator: Mathias E. Engels
creator: Eric C. Smidt
creator: Celice A. Silva
creator: Ana Kelly Koch
rights: Copyright (c) 2024 Lankester Botanical Garden, University of Costa Rica https://creativecommons.org/licenses/by-nc-nd/3.0/cr/
date: 2026-05-06
publication date: 2026-05-06
starting page: 222
ending page: 231
doi: 10.15517/lank.v24i3.61741

new green-flowered Aa (Cranichideae) from Peru
type: item
description: A new species from the Peruvian Andes, Aa olivacea, is described and illustrated. Information concerning distribution, habitat, ecology, and conservation status is provided. The new species is similar to A. hieronymi by its dense spikes of dark green and brown flowers with translucent whitish floral bracts; but differs from it by its elliptic to lanceolate leaves, 60 to 100-flowered spikes, glabrous rachis, lateral sepals with the margin entire to occasionally minutely erose near the apex, petals with slightly erose to sinuate apical margin, lip unlobed with lacerate to erose margin, and ovary with scarce hairs. Furthermore, A. tenebrosa is proposed as a synonym of A. hieronymi, A. nigrescens is recognized as a distinct species from A. leucantha, and A. lehmannii is confirmed to be an illegitimate, superfluous name for A. leucantha.
subject: Articles
  topic: section
subject: Aa hieronymi
  topic: keywords
subject: Aa leucantha
  topic: keywords
subject: Altensteinia
  topic: keywords
subject: Orchidaceae
  topic: keywords
subject: Peruvian Andes
  topic: keywords
subject: synonym
  topic: keywords
creator: Delsy Trujillo
creator: José D. Edquén
creator: Robin Fernandez-Hilario
creator: Akira A. Wong Sato
creator: Gerardo A. Salazar
rights: Copyright (c) 2024 Lankesteriana: International Journal on Orchidology https://creativecommons.org/licenses/by-nc-nd/3.0/cr/
date: 2026-05-06
publication date: 2026-05-06
starting page: 245
ending page: 257
doi: 10.15517/lank.v24i3.62472

new species Telipogon (Oncidiinae) from Bolivia
type: item
description: Telipogon minutus, from the Carrasco National Park in the Bolivian Yungas, is proposed as a new species. Plants of T. minutus, which reach up to 4 cm in height, are the smallest known Bolivian Telipogon. The new species is characterized by its oblong to ovate labellum, which is weakly sub-auriculate and retuse at the base, with a semi-circular stigma, a column with two lateral lobes and a mammillary protuberance at the apex, and two tufts of setae with trifurcate apices. A description, figures, and a distributional map of the new taxon are provided, along with a discussion of the differences between the new species and morphologically similar Telipogon species.
subject: Articles
  topic: section
subject: Andes
  topic: keywords
subject: Bolivian Yungas
  topic: keywords
subject: miniature Telipogon
  topic: keywords
subject: Orchidaceae
  topic: keywords
subject: taxonomy
  topic: keywords
creator: Modesto Zárate
creator: Carlos Martel
rights: Copyright (c) 2024 Lankesteriana: International Journal on Orchidology https://creativecommons.org/licenses/by-nc-nd/3.0/cr/
date: 2026-05-06
publication date: 2026-05-06
starting page: 233
ending page: 237
doi: 10.15517/lank.v24i3.62111

Rostellum in orchids
type: item
description: The rostellum, a small structure located above the stigma in the gynostemium of monandrous orchid flowers, separates the pollinia from the stigma, can prevent self-pollination, and plays an important role in pol- lination. It secretes a sticky exudate before and after anthesis, to which the pollinia are attached. Additionally,the viscidium, a sticky disk at the base of the stipe, also connects to the rostellum. This structure varies among species and holds taxonomic significance. For example, in Catasetum, the rostellum extends into antennae-like projections. After pollination, emasculation, or injury, the plant hormone ethylene is produced, triggering various post-pollination, post-emasculation, and post-injury processes, including changes in perianth color, senescence, and, ultimately, the death of floral segments that have completed their functions. Although the rostellum plays a significant role, it remains understudied, and many aspects of its function are still unknown. This review high- lights current knowledge and explores speculative hypotheses that may invite differing opinions and perspectives, which will likely be revised as future research deepens our understanding of its biological and ecological roles.
subject: Review
  topic: section
subject: emasculation
  topic: keywords
subject: ethylene
  topic: keywords
subject: floral senescence
  topic: keywords
subject: gynostemium
  topic: keywords
subject: pollinia
  topic: keywords
subject: pollination
  topic: keywords
subject: viscidium
  topic: keywords
creator: Hideka Kobayashi
creator: Joseph Arditti
rights: Copyright (c) 2024 Lankesteriana: International Journal on Orchidology https://creativecommons.org/licenses/by-nc-nd/3.0/cr/
date: 2026-05-06
publication date: 2026-05-06
starting page: 285
ending page: 318
doi: 10.15517/lank.v24i3.63156

Ten new orchid records from the Indonesian archipelago
type: item
description: Orchidaceae is one of the most diverse plant families in Indonesia, yet many species remain poorly documented across the archipelago. Here we report ten orchid species as new national records for Indonesia: Bulbophyllum nematocaulon, Bulbophyllum sanguineomaculatum, Cleisomeria lanatum, Corybas calopeplos, and Corybas holttumii from Sumatra; Acanthophippium bicolor and Anoectochilus papuanus from Java; Dendrobium teretifolium from the Lesser Sunda Islands, Bulbophyllum thiurum from Kalimantan; and Aerides agustiana from Sulawesi. For each species, we provide taxonomic notes, distributional data, type information, phenology, ecological observations, line drawings, and photographs. These new records extend the known geographic ranges of the species and confirm the importance of field exploration and herbarium study for documenting Indonesia’s remarkable orchid diversity.
subject: Articles
  topic: section
subject: epiphyte
  topic: keywords
subject: terrestrial
  topic: keywords
subject: Sundaland
  topic: keywords
subject: Lesser Sunda
  topic: keywords
subject: Orchidaceae
  topic: keywords
creator: Yuda R. Yudistira
creator: Aninda R. U. Wibowo
creator: Wendy A. Mustaqim
creator: Hedy Soleman
creator: Toni Artaka
creator: Richa K. Wati
rights: Copyright (c) 2026 Lankesteriana: International Journal on Orchidology https://creativecommons.org/licenses/by-nc-nd/3.0/cr/
date: 2026-04-24
publication date: 2026-04-24
starting page: 27
ending page: 48
doi: 10.15517/8g9n2n82

Orchids naturalized in Florida
type: item
description: Florida’s naturalized orchids are mostly terrestrials (10/15, 5/10 epiphytes), horticultural escapes (14/15), native to Asia (10/15), tropical American (4/10), and Africa (1/15). Only 3 species are widespread in Florida, Eulophia graminea, Eulophia maculata, and Zeuxine strateumatica, and can be considered invasive based on their spread beyond the areas of their initial establishments. Most of the other orchid species occur in one or two counties and have spread little so are considered naturalized but not invasive. The longest naturalized orchids (5/15) escaped more than 50 years ago, another group (5/10) more than 20 years ago, while the most recently naturalized species (5/10) less than 20 years ago. The 15 naturalized orchids comprise 12% (15/117 species) of Florida’s orchid flora, which is higher than most other places except for Hawaii’s (18/21) which has only 3 native orchids. Most orchids naturalized in Florida (10/15) have also naturalized elsewhere in the world. Failure to reproduce is the reason why most orchids don’t naturalize. To attempt to understand how Florida’s 15 naturalized orchids have overcome this barrier, the literature was examined to learn what their breeding systems and pollinators are in their native areas. This enabled predictions to be made about how these orchids reproduce in Florida. Four of the orchids need no pollinators because they are autogamous or apomictic. The native region pollinators of two of the orchids occur in Florida, five of the orchids are probably pollinated by congeneric species of their native area pollinators (Apis, Centris and Xylocopa), Two of the orchids are pollinated by close analogues of their native area pollinators (butterflies and hummingbirds). Overcoming other barriers to their naturalization, such as the lack of suitable mycorrhizal fungi or the inability to deal with abiotic limitations, has occurred but are poorly known and understood.
subject: Articles
  topic: section
subject: Eulophia
  topic: keywords
subject: horticulture
  topic: keywords
subject: naturalization barriers
  topic: keywords
subject: orchid floras
  topic: keywords
subject: ornamental trade
  topic: keywords
subject: pollinators
  topic: keywords
subject: Zeuxine
  topic: keywords
creator: Robert W. Pemberton
rights: Copyright (c) 2026 Lankesteriana: International Journal on Orchidology https://creativecommons.org/licenses/by-nc-nd/3.0/cr/
date: 2026-04-30
publication date: 2026-04-30
starting page: 65
ending page: 76
doi: 10.15517/sjs8k094

The genus Encyclia Hook. (Orchidaceae, Epidendroideae) in Maranhão state, Brazil: new morphological data and extent of distribution
type: item
description: In this study, we report the diversity, geographical distribution, and ecology of the Encyclia species from the state of Maranhão, Brazil. For each species treated, we present morphological descriptions, geographic distribution maps, taxonomic and ecological comments, an identification key for the species occurring in the state and photographic plates. Three species were recorded: E. gonzalezii, E. granitica, and E. linearifolioides, with E. gonzalezii being endemic to Brazil. Encyclia granitica has been locally rediscovered approximately 93 years after its only known collection in Maranhão. We also included new morphological data for this species, for which a description of the fruits had not been available until now. We found that the epicarp of the three species exhibited significant morphological differences, which may aid in their identification and delimitation. Given the complicated taxonomy of the genus, further studies focused on the epicarp may determine the extent to which it can assist in distinguishing additional species and in assessing its phylogenetic signal.
subject: Articles
  topic: section
subject: Amazon Forest
  topic: keywords
subject: Cerrado
  topic: keywords
subject: fruits
  topic: keywords
subject: orchids
  topic: keywords
subject: taxonomy
  topic: keywords
creator: Jaireslane F. Ribeiro
creator: Miguel Sena de Oliveira
creator: Maycon J. C. Silva
creator: Mizalene S. Silva
creator: Ellen T. S. Castro
creator: Amanda Leticia Garcia
creator: Lucas C. Marinho
creator: Alessandro W. C. Ferreira
rights: Copyright (c) 2026 Lankesteriana: International Journal on Orchidology https://creativecommons.org/licenses/by-nc-nd/3.0/cr/
date: 2026-04-27
publication date: 2026-04-27
starting page: 49
ending page: 64
doi: 10.15517/1edaz517

More than a century later: first modern record of Pteroglossa macrantha (Orchidaceae: Spiranthinae) in Argentina, with notes on its morphology, distribution, habitat, and conservation implications
type: item
description: Abstract. Pteroglossa macrantha (Orchidaceae: Spiranthinae) was previously known in Argentina only from a historical collection made by Ekman in 1908 in the province of Misiones, with no subsequent records confirming its presence in the country. During recent botanical surveys in the province of Corrientes, a new population was discovered, representing the first modern record for Argentina after more than a century and one of the southernmost confirmed occurrences of the species in the country. Detailed morphological observations confirmed its identity and revealed slight variations in leaf proportions compared with specimens from Brazil and Paraguay. The plants of this species grow in seasonally flooded alluvial grasslands on sandy substrates near the Paraná River. The regional conservation assessment, based on IUCN criteria, indicates that P. macrantha should be considered Critically Endangered (CR) in Argentina due to its extremely small area of occupancy, its occurrence in a single locality, and ongoing habitat degradation. This finding highlights the importance of the riparian grasslands of Corrientes as biodiversity refuges and priority areas for the conservation of rare and threatened orchids of the Southern Cone.
subject: Articles
  topic: section
subject: Corrientes Province
  topic: keywords
subject: Paraná River
  topic: keywords
subject: riparian grasslands
  topic: keywords
subject: southern distribution limit
  topic: keywords
creator: Javier E. Florentín
creator: Nadia S. Delvescovo
creator: Diego L. Salariato
creator: Graciela Terada
creator: Héctor A. Keller
creator: Eduardo A. Flachsland
rights: Copyright (c) 2026 Lankesteriana: International Journal on Orchidology https://creativecommons.org/licenses/by-nc-nd/3.0/cr/
date: 2026-04-14
publication date: 2026-04-14
starting page: 1
ending page: 10
doi: 10.15517/jnh7g404

A new species of Pleurothallis (Pleurothallidinae) from the P. talpinaria complex, with notes on comparative floral morphology and pollination ecology
type: item
description: A new species of Pleurothallis subgenus Talpinaria, discovered in the department of Tolima, Central Andes of Colombia, is described, illustrated, and discussed. The new species, Pleurothallis vallejoi is compared with P. talpinaria from Colombia and Venezuela, P. trimeroglossa from Peru and Ecuador, P. jostii, presumed to occur in Ecuador, and P. gracilicolumna from Colombia. The new species is distinguished from the previous ones by a lip with subquadrate lateral lobes, with narrowly elliptical basal auricles, and a callus that is elongated, thin, and slightly elevated, with the median lobe occupying almost half of the total lip length. Additionally, the pollination mechanism of the P. talpinaria complex is described and discussed.
subject: Articles
  topic: section
subject: Andean orchids
  topic: keywords
subject: Diptera
  topic: keywords
subject: Pleurothallis gracilicolumna
  topic: keywords
subject: Pleurothallis jostii
  topic: keywords
subject: Pleurothallis talpinaria
  topic: keywords
subject: Pleurothallis trimeroglossa
  topic: keywords
subject: Sylvicola
  topic: keywords
creator: Mario Alexei Sierra-Ariza
creator: José Armando Alvarez-Diaz
creator: Fernando Tinoco
creator: Mark Wilson
rights: Copyright (c) 2025 Lankesteriana: International Journal on Orchidology https://creativecommons.org/licenses/by-nc-nd/3.0/cr/
date: 2025-10-15
publication date: 2025-10-15
starting page: 151
ending page: 166
doi: 10.15517/zzdx7h51

Addition of Five New Species to the Orchid Flora of Bhutan IV
type: item
description: Five orchid species belonging to four genera, namely, Bulbophyllum deergongense, Bulbophyllum nepalense, Liparis sootenzanensis, Spiranthes himalayensis and Zeuxine gracilis, are reported for the first time from Bhutan. These findings are based on systematic native floristic diversity field surveys carried out between 2023 and 2025. We provide descriptions, along with information on type, distribution, phenology, habitat, notes, and colour plates to facilitate accurate species identification and support practical conservation efforts.
subject: Articles
  topic: section
subject: Orchidaceae
  topic: keywords
subject: section Brachyantha
  topic: keywords
subject: section Reptantia
  topic: keywords
subject: taxonomy
  topic: keywords
subject: Biodiversity
  topic: keywords
creator: Phub Gyeltshen
creator: Kinley Rabgay
creator: Karma Sangay
creator: Pema Tobgay
creator: Kezang Tobgay
creator: Pankaj Kumar
creator: Sangay Dema
rights: Copyright (c) 2026 Lankesteriana: International Journal on Orchidology https://creativecommons.org/licenses/by-nc-nd/3.0/cr/
date: 2026-04-21
publication date: 2026-04-21
starting page: 11
ending page: 25
doi: 10.15517/nxv7dt58